Eadweard Muybridge (1830-1904), US photographer and inventor, showing his instananeous photographs of animals in motion at the Royal Society. Muybridge invented a shutter which allowed an exposure of 1/500 seconds and using a battery of between 12 and 24 cameras, was able in 1877 to show that a trotting horse had all of its feet off the ground at times. In 1884-5, he carried out an extensive survey of the ovements of animals and humans for the University of Pennsylvania, publishing the results as 'Animal Locomotion' (1887).

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