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SMAD4 protein domain bound to DNA, molecular model. This strand of DNA (deoxyribonucleic acid, red and blue) is surrounded by MH1 domains of the SMAD4 (Mothers against decapentaplegic homolog 4) protein. This protein is forming a dimeric complex with the DNA. SMAD4 is involved in cell signalling, and when bound to DNA acts as a transcription factor.

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